Tables
1 Review of the worlds Haliplidae, the faunal regions where they occur 35
Figures
1, 2 Faunal regions 4
3–7 Immature stages of Haliplidae 7
8 Cladogram of Haliplidae 11
9 Haliplus flavicollis 25
10 Haliplus flavicollis 28
11–16 Haliplus spec. 29
17–19 Haliplus flavicollis 30
20–24 Haliplus flavicollis 31
Plates
1 Brychius elevatus 173
2 Brychius glabratus 174
3 Brychius hornii 175
4 Brychius hungerfordi 176
5 Brychius pacificus 177
6 Haliplus (Algophilus) lathridioides 178
7 Haliplus (Apteraliplus) parvulus 179
8 Haliplus (Australiplus) sindus 180
9 Haliplus (Borealiplus) borealis 181
10 Haliplus (Borealiplus) oblongus 182
11 Haliplus (Haliaplus) laminatus 183
12 Haliplus (Haliplidius) confinis 184
13 Haliplus (Haliplidius) obliquus 185
14 Haliplus (Haliplidius) rejseki 186
15 Haliplus (Haliplidius) subseriatus 187
16 Haliplus (Haliplidius) varius 188
17 Haliplus (Haliplus) aliae 189
18 Haliplus (Haliplus) apicalis 190
19 Haliplus (Haliplus) blanchardi 191
20 Haliplus (Haliplus) distinctus 192
21 Haliplus (Haliplus) dorsomaculatus 193
22 Haliplus (Haliplus) falli 194
23 Haliplus (Haliplus) fluviatilis 195
24 Haliplus (Haliplus) fulvicollis 196
25 Haliplus (Haliplus) furcatus 197
26 Haliplus (Haliplus) fuscicornis 198
27 Haliplus (Haliplus) harminae 199
28 Haliplus (Haliplus) heydeni 200
29 Haliplus (Haliplus) immaculatus 201
30 Haliplus (Haliplus) immaculicollis 202
31 Haliplus (Haliplus) interjectus 203
32 Haliplus (Haliplus) kamiyai 204
33 Haliplus (Haliplus) kirgisiensis 205
34 Haliplus (Haliplus) lineolatus 206
35 Haliplus (Haliplus) longulus 207
36 Haliplus (Haliplus) morii 208
37 Haliplus (Haliplus) ruficollis 209
38 Haliplus (Haliplus) samojedorum 210
39 Haliplus (Haliplus) sibiricus 211
40 Haliplus (Haliplus) simplex 212
41 Haliplus (Haliplus) stagninus 213
42 Haliplus (Haliplus) steppensis 214
43 Haliplus (Haliplus) turkmenicus 215
44 Haliplus (Haliplus) zacharenkoi 216
45 Haliplus (Liaphlus) abbreviatus 217
46 Haliplus (Liaphlus) africanus 218
47 Haliplus (Liaphlus) alastairi 219
48 Haliplus (Liaphlus) alluaudi 220
49 Haliplus (Liaphlus) andalusicus 221
50 Haliplus (Liaphlus) angusi 222
51 Haliplus (Liaphlus) angustifrons 223
52 Haliplus (Liaphlus) annulatus 224
53 Haliplus (Liaphlus) arrowi 225
54 Haliplus (Liaphlus) astrakhanus 226
55 Haliplus (Liaphlus) australis 227
56 Haliplus (Liaphlus) bachmanni 228
57 Haliplus (Liaphlus) basinotatus 229
58 Haliplus (Liaphlus) bonariensis 230
59 Haliplus (Liaphlus) brasiliensis 231
60 Haliplus (Liaphlus) camposi 232
61 Haliplus (Liaphlus) carinatus 233
62 Haliplus (Liaphlus) chinensis 234
63 Haliplus (Liaphlus) colombiensis 235
64 Haliplus (Liaphlus) columbiensis 236
65 Haliplus (Liaphlus) concolor 237
66 Haliplus (Liaphlus) confluentus 238
67 Haliplus (Liaphlus) costaricanus 239
68 Haliplus (Liaphlus) coxaspinipes 240
69 Haliplus (Liaphlus) crassus 241
70 Haliplus (Liaphlus) cribrarius 242
71 Haliplus (Liaphlus) cubensis 243
72 Haliplus (Liaphlus) cylindricus 244
73 Haliplus (Liaphlus) dalmatinus 245
74 Haliplus (Liaphlus) diopus 246
75 Haliplus (Liaphlus) discessus 247
76 Haliplus (Liaphlus) drechseli 248
77 Haliplus (Liaphlus) ebolovensis 249
78 Haliplus (Liaphlus) elsaltous 250
79 Haliplus (Liaphlus) excoffieri 251
80 Haliplus (Liaphlus) eximius 252
81 Haliplus (Liaphlus) exsecratus 253
82 Haliplus (Liaphlus) ferruginipes 254
83 Haliplus (Liaphlus) figuratus 255
84 Haliplus (Liaphlus) fiorentini 256
85 Haliplus (Liaphlus) flavicollis 257
86 Haliplus (Liaphlus) fulvus 258
87 Haliplus (Liaphlus) fuscipennis 259
88 Haliplus (Liaphlus) gamma 260
89 Haliplus (Liaphlus) garambanus 261
90 Haliplus (Liaphlus) genieri 262
91 Haliplus (Liaphlus) gracilis 263
92 Haliplus (Liaphlus) gravidoides 264
93 Haliplus (Liaphlus) gravidus 265
94 Haliplus (Liaphlus) guignoti 266
95 Haliplus (Liaphlus) guttatus 267
96 Haliplus (Liaphlus) halsei 268
97 Haliplus (Liaphlus) havaniensis 269
98 Haliplus (Liaphlus) heppneri 270
99 Haliplus (Liaphlus) incrassatus 271
100 Haliplus (Liaphlus) indicus 272
101 Haliplus (Liaphlus) indistinctus 273
102 Haliplus (Liaphlus) insularis 274
103 Haliplus (Liaphlus) javanicus 275
104 Haliplus (Liaphlus) kantneri 276
105 Haliplus (Liaphlus) kapuri 277
106 Haliplus (Liaphlus) kotoshonis 278
107 Haliplus (Liaphlus) lamottei 279
108 Haliplus (Liaphlus) langleyi 280
109 Haliplus (Liaphlus) leechi 281
110 Haliplus (Liaphlus) lewisii 282
111 Haliplus (Liaphlus) maculatus 283
112 Haliplus (Liaphlus) maculicollis 284
113 Haliplus (Liaphlus) maculipennis 285
114 Haliplus (Liaphlus) madagascariensis 286
115 Haliplus (Liaphlus) megapunctatus 287
116 Haliplus (Liaphlus) mesoamericanus 288
117 Haliplus (Liaphlus) methneri 289
118 Haliplus (Liaphlus) mexicanus 290
119 Haliplus (Liaphlus) mimulus 291
120 Haliplus (Liaphlus) minimus 292
121 Haliplus (Liaphlus) mucronatus 293
122 Haliplus (Liaphlus) nanus 294
123 Haliplus (Liaphlus) napolovi 295
124 Haliplus (Liaphlus) natalensis 296
125 Haliplus (Liaphlus) nedungaduensis 297
126 Haliplus (Liaphlus) nieseri 298
127 Haliplus (Liaphlus) nigerianus 299
128 Haliplus (Liaphlus) nitens 300
129 Haliplus (Liaphlus) novateutonianus 301
130 Haliplus (Liaphlus) oklahomensis 302
131 Haliplus (Liaphlus) ornatipennis 303
132 Haliplus (Liaphlus) ovalis 304
133 Haliplus (Liaphlus) panamanus 305
134 Haliplus (Liaphlus) perroti 306
135 Haliplus (Liaphlus) peruanus 307
136 Haliplus (Liaphlus) philippinus 308
137 Haliplus (Liaphlus) pilbaraensis 309
138 Haliplus (Liaphlus) pruthii 310
139 Haliplus (Liaphlus) pulchellus 311
140 Haliplus (Liaphlus) regili. 312
141 Haliplus (Liaphlus) rubidus 313
142 Haliplus (Liaphlus) rufescens 314
143 Haliplus (Liaphlus) samosirensis 315
144 Haliplus (Liaphlus) sharpi 316
145 Haliplus (Liaphlus) signatipennis 317
146 Haliplus (Liaphlus) signatus 318
147 Haliplus (Liaphlus) solitarius 319
148 Haliplus (Liaphlus) srilankanus 320
149 Haliplus (Liaphlus) stepheni 321
150 Haliplus (Liaphlus) sulawesicus 322
151 Haliplus (Liaphlus) tantoyucanus 323
152 Haliplus (Liaphlus) tanzanianus 324
153 Haliplus (Liaphlus) testaceus 325
154 Haliplus (Liaphlus) testudo 326
155 Haliplus (Liaphlus) thoracicus 327
156 Haliplus (Liaphlus) timmsi 328
157 Haliplus (Liaphlus) tocumenus 329
158 Haliplus (Liaphlus) triplehorni 330
159 Haliplus (Liaphlus) tumidus 331
160 Haliplus (Liaphlus) unicarinatus 332
161 Haliplus (Liaphlus) uniformis 333
162 Haliplus (Liaphlus) valdiviensis 334
163 Haliplus (Liaphlus) vancouverensis 335
164 Haliplus (Liaphlus) varicator 336
165 Haliplus (Liaphlus) variegatus 337
166 Haliplus (Liaphlus) venustus 338
167 Haliplus (Liaphlus) volpei 339
168 Haliplus (Liaphlus) watsoni 340
169 Haliplus (Liaphlus) wattsi 341
170 Haliplus (Liaphlus) youngi 342
171 Haliplus (Neohaliplus) lineatocollis 343
172 Haliplus (Neohaliplus) ruficeps 344
173 Haliplus (Nipponiplus) japonicus 345
174 Haliplus (Nipponiplus) latreillei 346
175 Haliplus (Nipponiplus) regimbarti 347
176 Haliplus (Paraliaphlus) alberti 348
177 Haliplus (Paraliaphlus) apostolicus 349
178 Haliplus (Paraliaphlus) connexus 350
179 Haliplus (Paraliaphlus) deceptus 351
180 Haliplus (Paraliaphlus) fasciatus 352
181 Haliplus (Paraliaphlus) grandis 353
182 Haliplus (Paraliaphlus) kenneri 354
183 Haliplus (Paraliaphlus) leopardus 355
184 Haliplus (Paraliaphlus) lucigelinae 356
185 Haliplus (Paraliaphlus) mimeticus 357
186 Haliplus (Paraliaphlus) mutchleri 358
187 Haliplus (Paraliaphlus) pantherinus 359
188 Haliplus (Paraliaphlus) pseudofasciatus 360
189 Haliplus (Paraliaphlus) punctatus 361
190 Haliplus (Paraliaphlus) tortilipenis 362
191 Haliplus (Paraliaphlus) triopsis 363
192 Peltodytes (Neopeltodytes) bradleyi 364
193 Peltodytes (Neopeltodytes) callosus 365
194 Peltodytes (Neopeltodytes) darlingtoni 366
195 Peltodytes (Neopeltodytes) dietrichi 367
196 Peltodytes (Neopeltodytes) dunavani 368
197 Peltodytes (Neopeltodytes) duodecimpunctatus 369
198 Peltodytes (Neopeltodytes) edentulus 370
199 Peltodytes (Neopeltodytes) festivus 371
200 Peltodytes (Neopeltodytes) floridensis 372
201 Peltodytes (Neopeltodytes) litoralis 373
202 Peltodytes (Neopeltodytes) muticus 374
203 Peltodytes (Neopeltodytes) oppositus 375
204 Peltodytes (Neopeltodytes) ovalis 376
205 Peltodytes (Neopeltodytes) pedunculatus 377
206 Peltodytes (Neopeltodytes) roughleyi 378
207 Peltodytes (Neopeltodytes) sexmaculatus 379
208 Peltodytes (Neopeltodytes) shermani 380
209 Peltodytes (Neopeltodytes) simplex 381
210 Peltodytes (Neopeltodytes) tamaulipensis 382
211 Peltodytes (Neopeltodytes) tortulosus 383
212 Peltodytes (Peltodytes) caesus 384
213 Peltodytes (Peltodytes) congoensis 385
214 Peltodytes (Peltodytes) coomani 386
215 Peltodytes (Peltodytes) dauricus 387
216 Peltodytes (Peltodytes) ghanaensis 388
217 Peltodytes (Peltodytes) intermedius 389
218 Peltodytes (Peltodytes) nodieri 390
219 Peltodytes (Peltodytes) pekinensis 391
220 Peltodytes (Peltodytes) quadratus 392
221 Peltodytes (Peltodytes) rotundatus 393
222 Peltodytes (Peltodytes) sinensis 394
223 Peltodytes (Peltodytes) speratus 395
224 Peltodytes (Peltodytes) sumatrensis 396
225 Peltodytes (Peltodytes) ugandanus 397
226 Phalilus bistriatus 398
227 Phalilus fortescueensis 399
228 Phalilus fuscatus 400
229 Phalilus gibbus 401
230 Phalilus hydei 402
231 Phalilus oberthuri 403
232 Phalilus pinderi 404
233 Phalilus storeyi 405
234 Photographs of Brychius elevatus – Haliplus (Borealiplus) borealis 409
235 Photographs of Haliplus (Borealiplus) oblongus – H. (Haliplus) apicalis 410
236 Photographs of Haliplus (Haliplus) blanchardi – H. (H.) harminae 411
237 Photographs of Haliplus (Haliplus) heyden – H. (H.) morii 412
238 Photographs of Haliplus (Haliplus) ruficollis – Haliplus (Liaphlus) abbreviatus 413
239 Photographs of Haliplus (Liaphlus) africanus – H. (L.) astrakhanus 414
240 Photographs of Haliplus (Liaphlus) australis – H. (L.) colombiensis 415
241 Photographs of Haliplus (Liaphlus) columbiensis – H. (L.) cylindricus 416
242 Photographs of Haliplus (Liaphlus) dalmatinus – H. (L.) exsecratus 417
243 Photographs of Haliplus (Liaphlus) ferruginipes – H. (L.) genieri 418
244 Photographs of Haliplus (Liaphlus) gracilis – H. (L.) incrassatus 419
245 Photographs of Haliplus (Liaphlus) indicus – H. (L.) langleyi 420
246 Photographs of Haliplus (Liaphlus) leechi – H. (L.) methneri 421
247 Photographs of Haliplus (Liaphlus) mexicanus – H. (L.) nieseri 422
248 Photographs of Haliplus (Liaphlus) nigerianus – H. (L.) peruanus 423
249 Photographs of Haliplus (Liaphlus) philippinus – H. (L.) sharpi 424
250 Photographs of Haliplus (Liaphlus) signatipennis – H. (L.) testaceus 425
251 Photographs of Haliplus (Liaphlus) testudo – H. (L.) valdiviensis 426
252 Photographs of Haliplus (Liaphlus) vancouverensis – Haliplus (Neohaliplus) lineatocollis 427
253 Photographs of Haliplus (Neohaliplus) ruficeps – H. (Paraliaphlus.) fasciatus 428
254 Photographs of Haliplus (Paraliaphlus) grandis – H. (P.) punctatus 429
255 Photographs of Haliplus (Paraliaphlus) tortilipenis – Peltodytes. (Neopeltodytes) edentulus 430
256 Photographs of Peltodytes (Neopeltodytes) festivus – P. (N.) sexmaculatus 431
257 Photographs of Peltodytes (Neopeltodytes) shermani – Peltodytes (Peltodytes) ghanaensis 432
258 Photographs of Peltodytes (Peltodytes) intermedius – P. (P.) ugandanus 433
259 Photographs of Phalilus bistriatus – P. storey 434
260 Distribution of Nearctic Haliplidae 437
261 Distribution of Nearctic Haliplidae 438
262 Distribution of Nearctic and Neotropical Haliplidae 439
263 Distribution of Nearctic and Neotropical Haliplidae 440
264 Distribution of Nearctic and Neotropical Haliplidae 441
265 Distribution of Nearctic and Neotropical Haliplidae 442
266 Distribution of Neotropical Haliplidae 443
267 Distribution of Neotropical Haliplidae 444
268 Distribution of Neotropical and Palaearctic Haliplidae 445
269 Distribution of Nearctic and Palaearctic Haliplidae 446
270 Distribution of Palaearctic Haliplidae 447
271 Distribution of Palaearctic Haliplidae 448
272 Distribution of Palaearctic Haliplidae 449
273 Distribution of Nearctic, Palaearctic and Oriental Haliplidae 450
274 Distribution of Palaearctic Haliplidae 451
275 Distribution of Palaearctic and Oriental Haliplidae 452
276 Distribution of Oriental and Australasian Haliplidae 453
277 Distribution of Australasian Haliplidae 454
278 Distribution of Australasian and Afrotropical Haliplidae 455
279 Distribution of Afrotropical Haliplidae 456
280 Distribution of Afrotropical Haliplidae 457
281 Distribution of Afrotropical Haliplidae 458